The Genus Euglena Ehrenberg 1838

(Authorities Note: The genus was first named in 1830, but was not diagnosed until 1838.)

Ehrenberg, C. G. 1830. Neue Beobachtungen uber blutartige Ercheinungnenin Aegypten, Arabien, und Sibirien, nebst einer Uebersicht und Kritik der fruher bekannten. Annalen der Physik 18:477-514.

Ehrenberg, C. G. 1838. Die Infusionsthierchen als vollkommene Organismen. Leipzig:Verlag von Leopold Voss.


Diagnosis:

Green, phototrophic chloroplast containing cells. The chloroplasts may be discoid, shield-shaped or ribbon-shaped, entire or dissected, with or without pyrenoids. Pyrenoids may be naked,or sheathed with one or two (double-sheathed) paramylon grains. Each chloroplast type is characteristic of a particular group of species. An eyespot is present at the anterior of the cell independent of the chloroplast. A paraflagellar body is present on the portion of the flagellum located in the reservoir. The flagellum emerges from the canal/reservoir complex subapically.The paraflagellar body is positioned adjacent to the eyespot. Cells are never completely rigid, but show some level of euglenoid movement (metaboly) ranging from slight to extreme.The cell body is asymmetrical and may be somewhat flattened. Cells usually have a single emergent flagellum, but in some species the flagellum is so short that it is not detectable with the light microscope. A second non-emergent flagellum is present in the reservoir. Never more than one active flagellum is present. A contractile vacuole is present in both marine and freshwater forms and expels into the reservoir. There tends to be much variation in shape, metaboly, chloroplasts and pyrenoids, and flagellar length among the species making identification difficult and many descriptions suspect.